The forest is a rich mix of temperate and tropical elements. Licuala palms, epiphytic orchids, and plants in the spurge (Euphorbiaceae) and shrimp plant (Acanthaceae) families have tropical origins, while maple, yew, aspidistra, and Paris are typically temperate. Multiple sets of specimens are collected for herbaria at the Russian Academy of Sciences, the Institute of Ecology and Bioresources, and the Missouri Botanical Gardens.